Villamarín-Cortez, Santiago. “Dung Beetles (Coleoptera: Scarabaeinae) of El Goaltal, Carchi, Ecuador: Annotated List of Species and Ecology”. ACI Avances En Ciencias E Ingenierías, vol. 2, no. 3, Dec. 2010, https://doi.org/10.18272/aci.v2i3.52.