M. H. Yánez-Muñoz, D. A. Veintimilla, E. N. Smith, M. Altamirano-Benavides, and C. J. Franklin, “Discovery of two surviving populations of Harlequin Toads (Amphibia: Bufonidae: Atelopus) in Ecuadorian Andes”, ACI Av. Cienc. Ing. (Quito), vol. 2, no. 3, Dec. 2010, doi: 10.18272/aci.v2i3.47.